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After trying times and bouts of melancholy, Jane finally publishes some novels, but illness soon looms. Cassandra tries to protect her sister's legacy, and her friend Isabella's future. 

Nancy's wedding to Roger is to take place in Poplar, but there's a surprise. Catherine prepares for her vows, and the midwives help a very young mother they have seen before.

When a boat nearly sinks due to sabotage, accusations fly around a fractured family. Are separated parents making a bid for time with their daughter? Is the grandmother trying to intervene?

Louis and Antoinette grow ever more threatened as the people begin to rise up and assert themselves against the monarchy.

While Cassandra spars with her sister-in-law Mary and tries to prepare a future for Isabella, she also thinks back to a long-ago romance, thanks to Jane's letters. 

Joyce faces her disciplinary hearing while the other midwives help an older mother who doubts her health and an younger struggling with an addiction.

As Christmas approaches, Pearl is asked to investigate a death by suicide even though most people doubt anything nefarious was involved.

As Antoinette grows even more unpopular and Louis begins to lose control, Antoinette accepts Lamballe back into her inner circle and tries to aid Louis in ruling.

After a death, Cassandra Austen rushes off to recover her sister Jane's letters, lest they reveal things to the public about the famous author and her family. 

Joyce tries to deal with an antagonistic patient but soon finds herself accused of negligence due to prejudice, while a garbage worker strike endangers Poplar's health. 

When a historical reenactment goes wrong, Mike and Nikki are brought in to investigate. Pearl, of course, eventually also finds herself involved – including, more and more, with Mike. 

Cardinal Rohan goes to trial and things quickly spin out of control, damaging Marie Antoinette's reputation as Louis' rivals scheme to take advantage. 

Cromwell hopes the king will grant him mercy as he is arrested and faces drummed up charges of treason. 

As Sister Julienne tries to help a woman with an abusive husband, she wonders if she's actually doing any good, while the other midwives aid a couple whose husband lives inside in an iron lung. 

Mike investigates the attempted murder of a real estate agent while Pearl looks into the sudden appearances of anonymous gifts to elderly people in town.